Assesses the macroevolutionary turnover of paleotropical planktic graptolites during the Late Ordovician Mass Extinction (LOME) via automated sequencing and capture-mark-recapture modeling. Graptolites exhibited a succession of turnover pulses (sensu Elizabeth Vrba) that were coincident with the main phases of the Hirnantian glaciation and during which the Diplograptina experienced declining metapopulation size, elevated extinction, zero species originations, and ultimately, complete exter...